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Ethan Carter, Ph.D.

Assistant Professor of Medicine



  Ethan Carter, Ph.D.
  Assistant Professor of Medicine


  University of Colorado Denver


Education & Training

University of Colorado, Boulder

1987 B.S. Kinesiology
1989 M.S. Exercise Physiology

University of Minnesota
1995 Ph.D. Physiology & Biophysics

University of California, San Francisco
1997 Post-Doc Physiology & Biophysics


Teaching Instructor, University of Colorado, Boulder

Research Assistant, Exercise Biochemistry Lab, Univ. of Colorado

Research Associate, Cardiology, Los Angeles Childrens’ Hospital

Research Assistant, Dept. of Physiology, Univ. of Minnesota

Post-doctoral research fellow, Cardiovascular Research Institute, UCSF

Instructor, Department of Physiology, UCSF

Post-doctoral research fellow, Cardiovascular-Pulmonary Research Lab, UCD

Assistant Professor, Dept. of Medicine, Univ. of Colorado Health Sciences Center

•Graduate School Fellowship, University of Minnesota (1990/91)
•Lifson-Johnson Memorial Award, Dept of Physiology, Univ. of Minnesota (1994)
•Proctor and Gamble Professional Research Award, American Physiological Society (1995)
•Fellow, American Lung Association (1996-98)

1.Carter, E. P. and M. A. Matthay. (1996). “Resolution of Pulmonary Edema - Recent Advances” in Current Pulmonology and Critical Care Medicine, Vol. 17: 351-376, Mosby-Year Book, Chicago IL.
2.Carter, E. P., C. Garat, T. Sakuma, Y. Wang, A. R. Campbell, and M. A. Matthay. (1998). “Resolution of alveolar edema: In vivo studies” in Pulmonary Edema. Vol. 116, Chap. 14, p. 431-455 of Lung Biology in Health and Disease series, Marcel Dekker, Inc. New York, NY.


1.Carter, E. P., S. E. Duvick, C. H. Wendt, J. Dunitz, L. Nici, O. D. Wangensteen, and D. H. Ingbar. (1994). Hyperoxia increases active alveolar Na+ resorption in vivo and in type II Na,K-ATPase in vitro. Chest. 105: 75S-78S.

2.Carter, E. P., M. A. Matthay, J. Farinas, and A. S. Verkman. (1996). Transalveolar osmotic and diffusional water permeability in intact mouse lung measured by a novel surface fluorescence method. J. Gen. Physiol. 108(3): 133-142. (See commentary by L. Reuss in same issue of J. Gen. Physiol.)

3.Umenishi, F., E. P. Carter, B. Yang, B. Oliver, M. A. Matthay, and A. S. Verkman. (1996). Sharp increase in rat lung water channel expression in the perinatal period. Am. J. Respir. Cell Mol. Biol. 15:673-679.

4.Carter, E. P., O. D. Wangensteen, S. O’Grady, and D. H. Ingbar. (1997). Effects of hyperoxia on type II cell Na,K-ATPase function and expression. Am. J. Physiol. 272 (Lung Cell. Mol. Physiol. 16):L542-L551.

5.Carter, E. P., F. Umenishi, M. A. Matthay, A. S. Verkman. (1997). Developmental changes in water permeability across the alveolar barrier in perinatal rabbit lung. J. Clin. Invest. 100:1071-1078.

6.Carter, E. P., O. D. Wangensteen, J. Dunitz, and D. H. Ingbar. (1997). Hyperoxic effects on alveolar sodium resorption and lung Na,K-ATPase. Am. J. Physiol. 273 (Lung Cell. Mol. Physiol. 17):L1191-L1202.

7.Lasnier, J. M., D. H. Ingbar, E. P. Carter, K. Wilson, S. McKnite, K. G. Lurie, and O. D. Wangensteen. (1998). Perfusion technique determines alveolar fluid resorption in the isolated perfused rat lung. J. Appl. Physiol. 84(2):740-745.

8.Dobbs, L., R. Gonzalez, M. A. Matthay, E. P. Carter, L. Allen and A. S. Verkman. (1998). Highly water-permeable type I alveolar epithelial cells confer high water permeability between the airspace and vasculature in rat lung. Proc. Natl. Acad. Sci. USA. 95(6):2991-2996.

9.Carter, E. P., B. P. Ölveczky, M. A. Matthay, and A. S. Verkman. (1998). High alveolar capillary endothelial water permeability in mouse lung measured by pleural surface fluorescence. Biophys. J. 74(4):2121-2128.

10.Garat, C., E. P. Carter, and M. A. Matthay. (1998). New in situ mouse model to quantify alveolar epithelial fluid clearance. J Appl. Physiol. 84(5):1763-1767.

11.Carter, E. P., K. Sato, Y. Morio, and I. F. McMurtry. (2000). Inhibition of KCa channels restores blunted hypoxic pulmonary vasoconstriction in rats with cirrhosis. Am. J. Physiol. Lung Cell. Mol. Physiol. 279:L903-L910.

12.Xu, L., E. P. Carter, M. Ohara, P-Y, Martin, B. Rogechev, K. Morris, M. A. Cadnapaphornchai, M. Knotek, and R. W. Schrier. (2000). Neuronal nitric oxide synthase and systemic vasodilation in rats with cirrhosis. Am. J. Physiol. Renal Physiol. 279:F1110-F1115.

13.Cadnapaphornchai M. A., M. Ohara, M. Knotek, B. Rogachev, T. Ladtkow, E. P. Carter, and R. W. Schrier. (2001). Role of nitric oxide in primary peripheral vasodilation, glomerular hyperfiltration, and increased renal plasma flow in normal pregnancy. Am. J. Physiol. Renal Physiol. 280:F592-F598.

14.Carter, E. P., C. L. Hartsfield, M. Jakkula, M. Miyazono, and I. F. McMurtry. (2001). Tissue specific regulation of heme oxygenase-1 during liver cirrhosis. In Press.

15.Miyazono, M., C. V. Garat, K. G Morris, Jr., and E. P. Carter. (2001). Decreased Renal Heme Oxygenase-1 Expression Contributes to Renal Vasoconstriction During Cirrhosis. In Press.

16.Carter, E. P., M. Miyazono, B. Fouty, and J. T. Reeves. (2001). New insights into the regulation of vascular tone during cirrhosis. In Preparation.